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Xorides humeralis (Say, 1828)

Anomalon humerale Say, 1828:74, ♀. ♀, Indiana; Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434, Neotype, type is lost (Academy Natural Science, Philadelphia); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:498, Type ♀ (destroyed); Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315, Holotype ♀, destroyed. - As Anomalon humeralis: - As Xorides humeralis: - Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474. (distribution); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:496 (comb. nov.); Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198. (distribution); Ruíz-Cancino et al., 2002Ruíz-Cancino, E., D.R. Kasparyan & J.M. Coronado Blanco 2002. Ichneumonidae. In: Llorente Bousquets, J. & J.J. Morrone (Ed.). Biodiversidad, taxonomía y biogeografía de artrópodos de México: Hacia una síntesis de su conocimiento, volumen III. 631-646. (distribution); Ruíz-Cancino et al., 2005Ruíz-Cancino, E. D.R. Kasparyan, S.N. Myartseva, V.A. Trjapitzin & J.M. Coronado Blanco 2005. Nuevos taxa de Hymenoptera Parasítica de Tamaulipas y otros estados de la República Mexicana. Biodiversidad Tamaulipeca 1:171-184. (distribution); Porter, 2005Porter, C.C. 2005. New records of Xorides (Hymenoptera: Ichneumonidae: Xoridinae) from Florida. Insecta Mundi 19 (3):187-190. (distribution); Chamé-Vázquez, 2019Chamé-Vázquez, E.R. 2019. Presencia de Xorides (Exomus) humeralis (Say, 1928) (Hymenoptera: Ichneumonidae: Xoridinae) en el estado de San Luis Potosí, México. Boletín de la Sociedad Entomológica Aragonesa 65:261-262. (distribution); Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319. (distribution). - As Xorides humeralis humeralis: - Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676. (distribution).
Xylonomus australis Cresson, 1870:167, ♀. ♀, Louisiana, Texas; Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434, Type (Acad. Nat. Sci. Philadelphia, No. 1520.1); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:498, Type ♀ (Quebec). - Cresson, 1870Cresson, E.T. 1870. Descriptions of new species belonging to the sub-family Pimplariae found in America north of Mexico. Transactions of the American Entomological Society 3:143-172. (distribution); Cresson, 1872Cresson, E.T. 1872. Hymenoptera Texana. Transactions of the American Entomological Society 4:153-292. (distribution); Dalla Torre, 1901-1902Dalla Torre, C.G. 1901-1902. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. — Volumen III: Trigonalidae, Megalyridae, Stephanidae, Ichneumonidae, Agriotypidae, Evaniidae, Pelecinidae. Pars II. Breitkopf & Härtel, Leipzig. VIII + 1141. (distribution); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:498 ( listed as synonym of Xorides humeralis humeralis (Say, 1828)); Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391 ( listed as synonym of Xorides humeralis (Say, 1828)); Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)). - As Xorides australis: - Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474. (distribution).
Xylonomus Lavallensis Provancher, 1874:59, Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434, Type ♀, labeled as humeralis with yellow label 517 in the 1877 Provancher collection (Public Museum, Quebec); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:498, Lectotype ♀ (Philadelphia). - Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:498 ( listed as synonym of Xorides humeralis humeralis (Say, 1828)); Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391 ( listed as synonym of Xorides humeralis (Say, 1828)); Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)). - As Xylonomus lavalensis: - Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434 ( listed as synonym of Xorides humeralis (Say, 1828)).
Xylonomus stigmapterus Say, 1904:pl. 9, fig. 2 - Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434 ( listed as synonym of Xorides piceatus (Rohwer, 1913)). - As Xorides stigmapterus: - Porter, 2005Porter, C.C. 2005. New records of Xorides (Hymenoptera: Ichneumonidae: Xoridinae) from Florida. Insecta Mundi 19 (3):187-190. (distribution).
Xylonomus piceatus Rohwer, 1913:357, Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.:434, Holotype (U.S.N.M., No. 15370); Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.:497, Type ♀ (Washington). Synonymized with Xorides humeralis (Say, 1828) by Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391. - Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)). - As Xorides humeralis piceatus: - Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676. (distribution), 497 (comb. nov.). - As Xorides piceatus: - Rohwer, 1920Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474. (distribution).
Xorides humeralis mexicanus Townes & Townes, 1960:497, ♀. Holotype ♀, 6 miles west of Antiguo, Mexico, 1,200 ft., July 21, 1954 (Lawrence, leg. University of Kansas Mexican Expedition). Synonymized with Xorides humeralis (Say, 1828) by Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391. - Townes & Townes, 1960Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676. (distribution); Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)).
Xorides humeralis sierrae Townes & Townes, 1960:500, ♂♀. Holotype ♀, Yosemite Park at 3,880 to 4,000 ft., June 31, 1931 (Berkeley). Synonymized with Xorides humeralis (Say, 1828) by Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391. - Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)).
Xorides humeralis excomptus Townes & Townes, 1960:500, ♂♀. Holotype ♀, Tanbark Flat, Los Angeles Co., Calif., June 25, 1956 (Davis, leg. R.C. Bechtel). Synonymized with Xorides humeralis (Say, 1828) by Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391. - Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)).
Xorides humeralis chiricensis Townes & Townes, 1960:501, ♀. Holotype ♀, Chiricahua Mts., Ariz., July 20, 1949 (Washington, USNM 63730, leg. Lindsay Kaiser). Synonymized with Xorides humeralis (Say, 1828) by Krombein et al., 1979Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.:391. - Khalaim et al., 2021Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.:315 ( listed as synonym of Xorides humeralis (Say, 1828)).

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Area distribution map

map of Xorides humeralis (Say, 1828)

Distribution

CANCanada
 (Chamé-Vázquez, 2019; Khalaim et al., 2021)
CAN-BrCCanada - British Columbia
 (Townes & Townes, 1960; Krombein et al., 1979)
CAN-ManCanada - Manitoba
 (Townes & Townes, 1960)
CAN-NeBCanada - New Brunswick
 (Townes & Townes, 1960)
CAN-NeFCanada - Newfoundland
 (Krombein et al., 1979)
CAN-OntCanada - Ontario
 (Rohwer, 1920; Townes & Townes, 1960)
CAN-QuéCanada - Québec
 (Rohwer, 1920; Townes & Townes, 1960 [as Xylonomus australis Cresson, 1870]; Townes & Townes, 1960)
MEXMexico
 (Townes & Townes, 1960 [as Xorides humeralis mexicanus Townes & Townes, 1960]; Ruíz-Cancino et al., 2002; Ruíz-Cancino et al., 2005)
MEX-NuLMexico - Nuevo León
 (Chamé-Vázquez, 2019; Khalaim et al., 2021)
MEX-SLPMexico - San Luis Potosi
 (Chamé-Vázquez, 2019 new; Khalaim et al., 2021)
MEX-TamMexico - Tamaulipas
 (Chamé-Vázquez, 2019; Khalaim et al., 2021)
USAUnited States
 (Chamé-Vázquez, 2019)
USA-AZUnited States - Arizona
 (Townes & Townes, 1960 [as Xorides humeralis chiricensis Townes & Townes, 1960]; Krombein et al., 1979)
USA-CAUnited States - California
 (Townes & Townes, 1960 [as Xorides humeralis sierrae Townes & Townes, 1960]; Townes & Townes, 1960 [as Xorides humeralis excomptus Townes & Townes, 1960]; Krombein et al., 1979)
USA-FLUnited States - Florida
 (Rohwer, 1920 [as Xorides piceatus (Rohwer, 1913)]; Rohwer, 1920 [as Xylonomus piceatus Rohwer, 1913]; Townes & Townes, 1960 [as Xylonomus piceatus Rohwer, 1913]; Townes & Townes, 1960 [as Xorides humeralis piceatus (Rohwer, 1913)]; Krombein et al., 1979; Porter, 2005 [as Xorides stigmapterus (Say, 1904)]; Porter, 2005)
USA-INUnited States - Indiana
 (Rohwer, 1920; Townes & Townes, 1960; Khalaim et al., 2021)
USA-KSUnited States - Kansas
 (Townes & Townes, 1960)
USA-LAUnited States - Louisiana
 (Cresson, 1870 [as Xylonomus australis Cresson, 1870]; Dalla Torre, 1901-1902 [as Xylonomus australis Cresson, 1870]; Rohwer, 1920 [as Xorides australis (Cresson, 1870)]; Townes & Townes, 1960 [as Xylonomus lavallensis Provancher, 1874])
USA-MAUnited States - Masachusettes
 (Rohwer, 1920; Townes & Townes, 1960)
USA-MDUnited States - Maryland
 (Rohwer, 1920; Townes & Townes, 1960)
USA-MIUnited States - Michigan
 (Townes & Townes, 1960)
USA-MNUnited States - Minnesota
 (Townes & Townes, 1960)
USA-MOUnited States - Missouri
 (Townes & Townes, 1960)
USA-MTUnited States - Montana
 (Townes & Townes, 1960)
USA-NCUnited States - North Carolina
 (Rohwer, 1920; Townes & Townes, 1960)
USA-NEUnited States - Nebraska
 (Townes & Townes, 1960)
USA-NHUnited States - New Hampshire
 (Townes & Townes, 1960)
USA-NJUnited States - New Jersey
 (Townes & Townes, 1960)
USA-NYUnited States - New York
 (Rohwer, 1920; Townes & Townes, 1960)
USA-OHUnited States - Ohio
 (Townes & Townes, 1960)
USA-ORUnited States - Oregon
 (Townes & Townes, 1960)
USA-PAUnited States - Pennsylvania
 (Townes & Townes, 1960)
USA-TXUnited States - Texas
 (Cresson, 1870 [as Xylonomus australis Cresson, 1870]; Cresson, 1872 [as Xylonomus australis Cresson, 1870]; Dalla Torre, 1901-1902 [as Xylonomus australis Cresson, 1870]; Rohwer, 1920; Rohwer, 1920 [as Xorides australis (Cresson, 1870)]; Townes & Townes, 1960 [as Xorides humeralis mexicanus Townes & Townes, 1960]; Townes & Townes, 1960; Krombein et al., 1979)
USA-UTUnited States - Utah
 (Townes & Townes, 1960)
USA-VAUnited States - Virginia
 (Rohwer, 1920; Townes & Townes, 1960)
USA-WIUnited States - Wisconsin
 (Townes & Townes, 1960)

Literature

  • Chamé-Vázquez, E.R. 2019. Presencia de Xorides (Exomus) humeralis (Say, 1928) (Hymenoptera: Ichneumonidae: Xoridinae) en el estado de San Luis Potosí, México. Boletín de la Sociedad Entomológica Aragonesa 65:261-262.
  • Cresson, E.T. 1870. Descriptions of new species belonging to the sub-family Pimplariae found in America north of Mexico. Transactions of the American Entomological Society 3:143-172.
  • Cresson, E.T. 1872. Hymenoptera Texana. Transactions of the American Entomological Society 4:153-292.
  • Dalla Torre, C.G. 1901-1902. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. — Volumen III: Trigonalidae, Megalyridae, Stephanidae, Ichneumonidae, Agriotypidae, Evaniidae, Pelecinidae. Pars II. Breitkopf & Härtel, Leipzig. VIII + 1141.
  • Khalaim, A.I., E. Ruíz-Cancino & J.M. Coronado-Blanco 2021. Darwin wasps of Mexico (Hymenoptera: Ichneumonidae): the subfamilies Acaenitinae, Poemeniinae and Xoridinae. Zoosystematica Rossica 30 (2):303-319.
  • Krombein, K.V., P.D. Hurd, D.R. Smith & B.D. Burks 1979. Catalog of Hymenoptera in America North of Mexico. Volume 1. Symphyta and Apocrita (Parasitica). Smithsonian Institution Press, Washington D.C. 1-1198.
  • Porter, C.C. 2005. New records of Xorides (Hymenoptera: Ichneumonidae: Xoridinae) from Florida. Insecta Mundi 19 (3):187-190.
  • Rohwer, S.A. 1920. The North American Ichneumon-flies of the tribes Labenini, Rhyssini, Xoridini, Odontomerini, and Phytodietini. Proceedings of the United States National Museum 57:405-474.
  • Ruíz-Cancino, E. D.R. Kasparyan, S.N. Myartseva, V.A. Trjapitzin & J.M. Coronado Blanco 2005. Nuevos taxa de Hymenoptera Parasítica de Tamaulipas y otros estados de la República Mexicana. Biodiversidad Tamaulipeca 1:171-184.
  • Ruíz-Cancino, E., D.R. Kasparyan & J.M. Coronado Blanco 2002. Ichneumonidae. In: Llorente Bousquets, J. & J.J. Morrone (Ed.). Biodiversidad, taxonomía y biogeografía de artrópodos de México: Hacia una síntesis de su conocimiento, volumen III. 631-646.
  • Say, T. 1828. A description of some new species of Hymenoptera of the United States. Contributions of the Maclurian Lyceum to the Arts and Sciences 1:67-83.
  • Townes, H.K. & M. Townes 1960. Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. Bulletin of the United States National Museum 216 (2):1-676.